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In Darwin’s Footsteps (Eastern Route) route map

Begin in Quito (9,350 ft) and explore its UNESCO-listed Old Town of colonial plazas and churches. Settle into your hotel for an overnight stay; optional pre-voyage extension available.

Guided tour of Quito’s Old Town with Ecuadorian chocolate tasting. Head 16 miles north to the ‘Middle of the World City’; after lunch, visit the Equator monument and stand with one foot in each hemisphere.

Meals: Lunch

After breakfast, fly to Baltra Island and transfer to board MS Santa Cruz II. Make a panga landing at Mosquera Islet to walk sandy dunes among sea lions, blue-footed boobies and Sally Lightfoot crabs with naturalist guides.

Meals: Breakfast

Ride by panga to Punta Pitt for a nature walk and the chance to see all three booby species, with options to explore by boat, snorkel or swim. Continue to the Cerro Colorado Tortoise Reserve to observe giant tortoises, San Cristóbal Mockingbirds and lava lizards.

On Santa Fé, take a nature walk on idyllic beaches to see endemic Barrington land iguanas and sea lions; optional swim/snorkel from panga, kayak or glass‑bottom boat. After lunch, visit South Plaza for frigatebirds, Swallow‑tailed Gulls, Galápagos Shearwaters, and land and marine iguanas.

Meals: Lunch

Disembark at Puerto Ayora to visit the Charles Darwin Research Station, with a giant tortoise breeding center, recently hatched tortoises, Darwin’s finches and a prickly pear cactus forest. Enjoy lunch in the highlands among wild tortoises, with time for another activity such as mountain biking or a sugar plantation visit.

Meals: Lunch

On Española, explore Punta Suárez for Nazca and blue-footed boobies, Swallow-tailed Gulls and (Apr–Dec) waved albatross; see red Española lava lizards, marine iguanas and the El Soplador blowhole. After lunch, continue to Gardner Bay and nearby Osborn Islet for sea lions, mockingbirds and finches, with optional snorkelling or kayaking.

Meals: Lunch

At Eden Islet, look for Galápagos green turtles, rays and reef sharks, and frigatebirds; options include swimming, snorkelling, glass-bottom boat or kayaking. After lunch, sail to North Seymour to see colonies of blue-footed boobies and Swallow-tailed Gulls, plus sea lions and marine iguanas.

Meals: Lunch

Disembark at Baltra Island and transfer to the airport in Baltra for your flight to Guayaquil or Quito to connect to your international flight; optional post-voyage extensions available.

What’s included

Meals served on board during the Galápagos cruise, prepared by the ship’s chefs.
Overnight hotel stay in Quito at the start of the trip.
Cabin accommodation aboard the expedition ship MS Santa Cruz II throughout the Galápagos sailing.
Transfers between Baltra Airport and the ship at embarkation and disembarkation.
Guided tour of Quito’s Old Town with a visit to the Equator’s ‘Middle of the World’ monument.
Guided shore excursions and nature walks at sites including Mosquera Islet, Punta Pitt, Santa Fé, South Plaza, Santa Cruz highlands, Española (Punta Suárez and Gardner Bay), Eden Islet, and North Seymour; opportunities to snorkel, kayak, swim, and ride the glass-bottom boat as offered by the expedition team.

What’s not included

International flights to and from Ecuador are not included.
Entry visas and travel documents (if required) are the traveller’s responsibility.
Travel insurance is not included and must be arranged by the traveller.
Meals not specified on land (e.g., in Quito).
Optional activities such as mountain biking or visiting a sugar plantation on Santa Cruz, unless expressly stated as included.
Personal expenses such as laundry, souvenirs, beverages, and phone calls.
Pre- and post-voyage extension options mentioned in the itinerary.

Additional information

Visit the famed Mitad del Mundo (Middle of the World City) north of Quito to stand astride the Equator. Pose at the yellow line and circle the 30-meter Equatorial Monument, where exhibits explain how explorers and scientists fixed the planet’s midpoint.

Engage with interactive displays on navigation, latitude, and hemispheres, and pause for photos and reflection at this geographical icon. The surrounding Andean views and open plazas add drama to a uniquely tangible global landmark.

Arrive by panga for a wet landing on sandy Mosquera Islet, a low-lying sandbar between Baltra and Seymour known for one of the archipelago’s largest Galápagos sea lion colonies. Walk the beach with your naturalist guide to observe relaxed harems and pups at close, respectful distances.

Scan tidepools for vivid Sally Lightfoot crabs and watch for blue-footed boobies plunge-diving just offshore. The flat, open terrain and bright volcanic sands create superb light for wildlife photography.

Walk the tuff-cone slopes of Punta Pitt, the only Galápagos site where you may see all three booby species—blue-footed, Nazca, and red-footed—along with frigatebirds riding the thermals. Trails reveal stratified ash landscapes with broad Pacific vistas.

Slip into clear water from the panga to snorkel among schools of reef fish, with chances for sea turtles and playful sea lions. Your guide interprets the island’s ecology, highlighting how currents and geology shape this rich bird habitat.

Visit the Cerro Colorado Tortoise Reserve, a conservation center safeguarding the archipelago’s emblematic reptiles. Walk shaded trails to see giant tortoises browsing in semi-wild enclosures designed to mirror their native highland forests.

Learn about head-start programs, nesting, and the species’ century-long lifespans. Keep an eye out for endemic companions like the San Cristóbal mockingbird and lava lizards, and hear how science and stewardship work together to protect fragile island ecosystems.

Walk powdery coves of Santa Fé Island in a turquoise bay frequented by sea lions. Trails weave among towering Opuntia cactus forests where the pale-hued, endemic Barrington land iguana often basks beside volcanic rocks.

Choose to snorkel over rocky reefs, kayak along calm shores, or ride a glass-bottom boat to spot rays and reef fish without getting wet. Guides share insights on island endemism and how arid-adapted flora sustains unique wildlife communities.

Follow a coastal path atop South Plaza Island, where sea cliffs host wheeling seabirds and the ground blazes with seasonal sesuvium “carpet” hues. Watch land iguanas lounge beneath prickly pear cacti, waiting for fallen pads and fruit.

Look for swallow-tailed gulls, shearwaters, and frigatebirds gliding on updrafts. The compact island concentrates wildlife encounters in dramatic scenery, offering excellent viewpoints for photography across cobalt channels and rust-red lava outcrops.

Visit the Charles Darwin Research Station in Puerto Ayora to see giant tortoise breeding pens, juvenile reptiles, and exhibits on evolution and conservation inspired by Darwin’s finches. Stroll through stands of prickly pear cacti frequented by land birds.

Continue into the lush Santa Cruz highlands for a cooler microclimate where wild giant tortoises graze and wallow. Over lunch and a relaxed walk, learn how rainfall, soils, and vegetation shape habitats across this volcanic island.

Hike the rugged trail at Punta Suárez, a sanctuary for Nazca and blue-footed boobies, marine iguanas, and—April to December—the waved albatross. Crashing surf and lava ledges create a powerful coastal stage for wildlife displays.

Pause at El Soplador, a natural blowhole that sends plumes skyward with each swell. Your guide explains nesting cycles, courtship rituals, and how isolated Española evolved distinct species and striking color morphs.

Relax on the white coral sands of Gardner Bay where curious sea lions doze along the shore and Galápagos mockingbirds flit between beach vines. Shallow, clear water makes this one of the archipelago’s most inviting swimming spots.

Snorkel near Osborn Islet to see reef fish, rays, and the occasional turtle, often with playful sea lions twisting through your bubbles. The broad bay and gentle swell create easy conditions for all experience levels.

Drift by panga to tranquil Eden Islet, watching for gliding rays and Galápagos green turtles in the shallows. On land, look for magnificent frigatebirds with inflatable scarlet throat pouches perched in low shrubs.

Choose to snorkel above rocky reefs alive with parrotfish and damselfish, kayak quiet coves, or view marine life from a glass-bottom boat. Guides interpret the nutrient-rich currents that power this productive coastal ecosystem.

Walk the level, uplifted lava of North Seymour through one of the Galápagos’ most active seabird breeding colonies. Witness blue-footed booby courtship and frigatebirds with scarlet gular sacs, while land iguanas and sea lions occupy the shoreline.

Interpretive stops reveal how tectonic uplift formed the island’s terrace and why nutrient-rich seas support dense wildlife. Low shrubs offer unobstructed viewing, ideal for photography of nests, displays, and coastal vistas.

Pedal quiet backroads in the Santa Cruz highlands, moving through misty Scalesia forests and pastoral clearings where tortoises sometimes graze. Rolling terrain and cool air make for an energizing ride with broad views toward the coast.

Guides tailor distance and pace, sharing insights on island microclimates, soils, and farming traditions. This experience is optional and bookable on tour at an additional cost, subject to local conditions and availability.

Step into a traditional trapiche (sugarcane mill) in the Santa Cruz highlands to see cane pressed into juice and cooked into panela. Taste small-batch spirits and sample locally grown cacao transformed into rich chocolate.

Hosts explain highland agriculture and family-run production that sustains island communities. This cultural experience is optional and bookable on tour at an additional cost, with timings coordinated around ship operations.

Explore Quito’s UNESCO-listed Historic Center with a local expert, pausing in elegant plazas and ornate churches around Plaza Grande and the baroque La Compañía de Jesús. Stroll cobblestone lanes lined with balconied mansions while your guide traces the story from Inca foundations to Spanish colonial Quito.

Sample Ecuador’s prized cacao with a guided tasting of artisan chocolate made from Arriba Nacional beans. Learn how altitude, terroir, and careful roasting shape rich aromas before time to admire Andean vistas and the Old Town’s tiled roofs.

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